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Massospondylus (/ˌmæsoʊˈspɒndɪləs/ MASS-oh-spon-di-lus[1]) is a genus of sauropodomorph dinosaur from the Early Jurassic of southern Africa. It was described by Richard Owen in 1854 from remains discovered in South Africa, and is thus one of the first dinosaurs to have been named. The name Massospondylus means 'longer vertebra', alluding to what Owen identified as tail vertebrae; these vertebrae are now known to be from the neck. Although the original fossils were destroyed in London during a bombing raid in World War II, a plethora of specimens have since been assigned to the genus, making it one of the best-known sauropodomorphs from the Early Jurassic. The genus lived during the Hettangian, Sinemurian, and Pliensbachian ages, which lasted from ca. 201 to 184 million years ago. Most fossils come from the upper Elliot and Clarens formations of South Africa and Lesotho, but the genus is also found in the Forest Sandstone and the Mpandi Formation of Zimbabwe. Material from the US, India, and Argentina was previously assigned to the genus, but the US and Argentinian specimens are now assigned to their own genera (Sarahsaurus and Adeopapposaurus). Because of their great abundance, Massospondylus fossils have been used to date rocks, and a biozone, the Massospondylus Range Zone, is named after the genus.
Two species are considered valid: the type species M. carinatus, as well as M. kaalae, which was named in 2009 and is known from a single skull. Six other species have been named during the past 150 years but are no longer recognised. Originally, Massospondylus and similar dinosaurs have been regarded as theropods, but are now classified as basal ("early diverging") members of Sauropodomorpha. This group also includes sauropods. Within sauropodomorphs, Massospondylus is often classified in the family Massospondylidae. The genus was 4–6 metres (13–20 ft) long, with a long neck and tail, a small head, and a slender body. It is distinguished from related genera by the very elongated vertebrae of the front portion of the neck, amongst other features. Although Massospondylus was long depicted as quadrupedal (four-legged), it is now considered to have been bipedal (two-legged).
It was probably a herbivore (plant-eater), although some have speculated that basal sauropodomorphs may have been omnivorous. On each of its hands, it bore a sharp thumb claw that was probably used in feeding, possibly to uproot vegetation or to pull down branches while rearing up. Clutches of eggs have been found, some of which contained embryos; these are among the oldest eggs and embryos of an amniote in the fossil record. The eggshell was extremely thin, less than 0.1 mm (0.0039 in), unlike the much thicker eggshells in later dinosaurs. The embryos had proportionally longer arms than adults and a very large head, leading researchers to suggest that they were quadrupedal and shifted to a bipedal posture later during growth. Newer research instead suggested that Massospondylus was bipedal at all ages. Individuals accelerated or slowed down their growth depending on environmental factors such as food availability. The oldest known specimen was around 20 years of age.
Two species are considered valid: the type species M. carinatus, as well as M. kaalae, which was named in 2009 and is known from a single skull. Six other species have been named during the past 150 years but are no longer recognised. Originally, Massospondylus and similar dinosaurs have been regarded as theropods, but are now classified as basal ("early diverging") members of Sauropodomorpha. This group also includes sauropods. Within sauropodomorphs, Massospondylus is often classified in the family Massospondylidae. The genus was 4–6 metres (13–20 ft) long, with a long neck and tail, a small head, and a slender body. It is distinguished from related genera by the very elongated vertebrae of the front portion of the neck, amongst other features. Although Massospondylus was long depicted as quadrupedal (four-legged), it is now considered to have been bipedal (two-legged).
It was probably a herbivore (plant-eater), although some have speculated that basal sauropodomorphs may have been omnivorous. On each of its hands, it bore a sharp thumb claw that was probably used in feeding, possibly to uproot vegetation or to pull down branches while rearing up. Clutches of eggs have been found, some of which contained embryos; these are among the oldest eggs and embryos of an amniote in the fossil record. The eggshell was extremely thin, less than 0.1 mm (0.0039 in), unlike the much thicker eggshells in later dinosaurs. The embryos had proportionally longer arms than adults and a very large head, leading researchers to suggest that they were quadrupedal and shifted to a bipedal posture later during growth. Newer research instead suggested that Massospondylus was bipedal at all ages. Individuals accelerated or slowed down their growth depending on environmental factors such as food availability. The oldest known specimen was around 20 years of age.
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